Clonal governance

Take any system where:

• units can self-replicate
• units mutate over time
• units form a higher-level cooperative body / swarm / superorganism
• selection pressures exist both inside the collective and at the collective level

“Maximum fitness” at the unit level selects for defection, over-replication, resource capture, and escape from specialization.
“Maximum cooperation” is a metastable attractor that has to be actively maintained by policing, bottlenecks, surveillance, differentiation, apoptosis/sacrifice norms, reproductive suppression, memory limits, spatial constraints, immune cleanup, etc.

The life or death question is:
How long can a clonal collective keep the maximum-cooperation replicator state from collapsing back toward the maximum-fitness replicator state?

So the analogical family for cellular senescence is broader than just multicellular