Here are “multi-polar traps” you keep running into when a lineage tries to become a Darwinian individual (an organism) while the old units (cells, nuclei, lineages) still have their own fitness gradients. “Multi-polar” here means there are multiple selection targets with partially incompatible objective functions: cells vs collective, different cell lineages vs each other, nuclei within a syncytium, and sometimes “fast” within-lifetime selection vs “slow” between-generation selection.

1. Relatedness collapse: aggregation and chimeras (the classic “different genotypes in one body” trap)
   If your multicellular stage is built by aggregation or frequent fusion, you get genetically mixed groups. Then within-group selection becomes loud: selfish lineages can