John Maynard Smith

John Maynard Smith (1920–2004) was a British evolutionary biologist and professor at the University of Sussex, known for applying game theory to evolutionary biology and for co-authoring the framework of major evolutionary transitions. Originally trained as an aeronautical engineer, he studied under J. B. S. Haldane before moving into genetics and evolutionary theory.
Evolutionary game theory
Maynard Smith introduced the concept of the evolutionarily stable strategy (ESS), a strategy that, when adopted by most members of a population, cannot be invaded by any rare alternative strategy. His book Evolution and the Theory of Games (1982) established contest behavior, sex ratios
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